Dispersal
editIt is probable that dispersal of Tarebia granifera from one waterbody or river catchment to another occurs passively via birds, notably waterfowl, which eat them and void them later, perhaps in another habitat.[1] Evidence for this comes from the finding of many small Tarebia granifera 5–7 mm in height and still containing the soft parts in unidentified bird droppings from the bank of the Mhlali River, South Africa.[1] Even though the shell of Tarebia granifera is thick, most of these juveniles had been partially crushed with only a few still intact.[1] Both the intact and damaged specimens could have been alive when passed and perhaps survived had they been deposited in water.[1] None was large enough to have been reproductively mature (see below) and would have needed to survive in any new habitat for several months before reproducing.[1]
Passive dispersal may also occur via weed on boats and boat trailers and via water pumped from one waterbody to another for industrial and irrigation purposes.[1] In the Nseleni River juvenile Tarebia granifera were commonly found with another invasive snail, Pseudosuccinea columella, on floating clumps of water hyacinth Eichhornia crassipes which provide a vehicle for rapid downstream dispersal.[1]
Once established in a particular waterbody Tarebia granifera is likely to disperse actively, both up and downstream in the case of flowing systems, as far as environmental factors like current speed and food availability will allow.[1] The snail's tolerance of turbulent, flowing water was demonstrated by Prentice (1983)[16] who reported it migrating upstream on the Caribbean island of Saint Lucia at a rate of 100 m month−1 in streams discharging up to 50 L.s−1.[1] In KwaZulu-Natal it has been collected in water flowing at up to 1.2 m.s−1 which is likely to exceed the current speeds of at least the lower and middle reaches of many rivers and streams in South Africa making these watercourses open to colonization.[1]
The sole of Tarebia granifera is proportionally small when compared to other thiarids and smaller snails with their higher coefficients were less able to grip the substratum in the face of moving water and so did not disperse as effectively as larger ones.[1]